very helpful. I have around 30 new sites done 2 monthes before penguin. I got around 100 backlinks (approved) from directory submission for each site in march and april just before penguin started. Most of these 30 sites ranked on page 3 to page 5 for the main target keywords since June this year. Couple of them even on page 1 or page 2. I didn't do any backlinking except those directory submissions done before penguin. So the directory submission did help. But all these new sites target long tail keywords, with search volumn under 10000 per month. (most are between 4000 to 8000 search per month)
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